Line art is the oldest way to make a picture: a line where a thing ends. Print taught it to carry tone as well. The engravers of the old floras shaded with hatching, laying lines closer where a leaf turned away from the light.
In 1949 Picasso drew in the air with a light for Gjon Mili’s camera. Hergé’s even contour in Tintin was later named ligne claire, the clear line, by Joost Swarte in 1977.
On screens it came back as outline icons. iOS 7 traded glossy pictures for thin lines in 2013, and most icon sets are still drawn on a 24-unit keyline grid. The rules have barely changed: one weight of line, closed shapes, and tone from lines laid side by side.